xt7cfx73wv2d https://exploreuk.uky.edu/dips/xt7cfx73wv2d/data/mets.xml Alabama Alabama Museum of Natural History 1948 Other titles include: Alabama Museum of Natural History museum paper, Geological Survey of Alabama, Museum of the Geological Survey of Alabama. Other creators include: United States. Work Projects Administration, Geological Survey of Alabama, Tennessee Valley Authority. Issues for 1, 3 carry no series numbering. No. 2 also as Education papers no. 1. UK holds archival copy for ASERL Collaborative Federal Depository Program libraries. Call number AS36 .A2. journals English University, Ala. : Alabama Museum of Natural History, 1910-1960 This digital resource may be freely searched and displayed in accordance with U. S. copyright laws. Alabama Works Progress Administration Publications Museum Paper, no. 27, 1948 - including "New Anophthalmid Beetles (Fam. Carabidae) from the Appalachian Region" by J. Manson Valentine text Museum Paper, no. 27, 1948 - including "New Anophthalmid Beetles (Fam. Carabidae) from the Appalachian Region" by J. Manson Valentine 1948 1948 2015 true xt7cfx73wv2d section xt7cfx73wv2d 3
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RJ GEOLOGICAL SURVEY OF ALABAMA
WALTER B. JONES, State Geologist 1
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MUSEUM PAPER 27 B
ALABAMA MUSEUM 0I= NATURAL HISTORY
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SEEE E
ml O, NEW ANOPHTHALMID BEETLES (FAM. CARABIDAE)
» FROM THE APPALACHIAN REGION
r by
` J. MANSON VALENTINE
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GEOLOGICAL SURVEY OF ALABAMA
WALTER B. JONES, State Geologist
MUSEUM PAPER 27
ALABAMA MUSEUM OF NATURAL HISTORY
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E ANOPHTHALMID BEETLES (FAM. CARABIDAE) ;
FROM THE ARPALAQHIAN REGION
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J. MANSON VALENTINE
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` INTRODUCTION
, n many years Dr. Valentine has carried out field and
s lrtht >ry studies of Anaphthalmid beetles, and has made im-
I port contributions to a fuller knowledge and understand-
Q ing hat group. It is intended that Dr. Valentine prepare l*
» a tw ctc paper on the group, to be published, along with
` siitti studies of other forms of cave faunas b other s ecial— 3
l ists. >ur lorthcoinmg monograph on caves and caverns ol
.»\lal a. However, many caves remain to be studied, so it
was » ded to publish certain specific data from time to time, §·
. . . . . . . F
V. that information might be available to all interested par-
T tics. ter on, the cave monograph will carry a complete dis-
cuss. »»l` the group.
Q .
t ACKNOIVLEDGEMENTS
I fl
L give our grateful thanks to jim Van Antwerp, Mo-
»_ btlt bama, who oenerousl t yrovided funds for Dublication if
. D . 5 l {
· ol tl tt ner. Also we wish to thank Harve Tem Jleton, Dr. tt
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I
I I V ANOPHTHALMID BEETLES (FAM. CARABIDAE)
_ FROM THE APPALACHIAN REGION
_}. MANSON VALENTINE
I I
I GENERAL DISCUSSION
. s
ace the tune ol publication ol the author’s last paper*”
I tleal _ with the anophthalinicl Iauna of the Appalachian
1 valli considerable material in the group has accumulated.
= Sun I the various collectings has revealed a nuinber of new
' sptt and geographic races at present referable to the genus
_ IM; .»oj;Izt/mImus Jeannel. These new Iorins are describ-
2 ell I ln.
- ze technique ol description Iollows very closely that
Q em] ed in the article relerred to above, which contains I
_ tlell >ns of nieasurenients and their derived indices as well
{ as ti itionary and distributional conclusions applicable to
‘ the sent work. In order better to interpret the data ap-
V pun below it is desirable to use the two papers in con-
I iunl 1.
I
breviations employed in description have the Iollow-
; ing tnings: L.—total length; \`V.—]lIZlXIlIIIllIl width: D.—
s mini;. nn depth: w.—width index: d.—depth index: h.——
j dpi I t index; p.—pronital index; e.»—elytral index; a.—an-
·‘ tw index: s.—antennal segmental index: c.—chaetotaxial
I W·l< g.—genital index.
. I
_ PW .u0pI1thaImus intermedius teinpleloni NOW 5Ul>$}>¢·
(`l
_ Q V/ze Ior·aIz"z‘_y: I‘IIggI1`1I)()[IIZllH’S Cave. McMinnville.
. ' I\’arren Co., Tenn.
2/»r’.t’ Qyrous, eonVCX. P
ll ‘/ { *7114: Narrow, convex, glabrous: humeri rounded,
l" Y `“l" *1flY arcuate, apex rounded; rellexed margin narrow. ,
Z" . Ml Gttate and ciliate at humeri, somewhat expanded api-
A (lull P()ll][ of juncture with epipleura; SLl`i21€ i21i1`ly WGH lIU·
" lm` V Yaguely punctured, anterior portion ol recurved
F Q1:
j .
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lll (Leo/og/czrl Survey of Alalzanm
apical stria relatively short, gently arcuate or parallel » su- i
ture: humeral papillae rather widely distributed, sprea river I
a distance nearly equal to three fourths that between 1 liu- l
meral and medial sets; humeral impressions shallow. tle- l
fined. disc only slightly flattened. .
;l[1]1em/ages: Jaws. palpi elongate, subtermin seg .
ment of maxillary palpi 6 t’7ths the length of terminal; aren-
nae. legs long, testaceous.
Amlmgztsz Median lobe long. narrow. cylindrica iasal
bulb well defined, basal flexure proximal; apical test ; M
short. narrow in dorsal profile. decurved; transfer ap `llllls ll"
relatively short, copulatory pieces two—an internal r antl s lull
an external enveloping piece, the latter spout—shaped rlor All
sal, broadly spatulate in laterial view; parameres el ;;1lt‘. Ul
with four long. stout setae. (Pl. ll, fig. 2) _
]
Four species of Pszjzzzlmiojilzllmlnius are now knov ana; ~ gn
l the Cumberland River drainage system. Althougli l tli· V lla
verse form (nmzrarlci and fIll("l`}Ht”(ll.ll5` elongate. rolms unl . ol
_ czznzlzcrlzzmlirs transverse). they show a certain kinslr in i —_ iw
general tendency toward convexity and in their coinpl lad; pri
of pubescence. a character rarely met with in the genus llel ‘·i· pa
is also considerable correspondence in aedeagel Ctlllf ~lt‘lll. g. mi
especially between the first three species mention dir ; in
ranges of which overlap in west central Tennessee. ( I ro
lzoztlzts inhabits the main river valley of the Cumberla _ an
It is Z1 great pleasure to name the fine species wltvl ·v its
t here in honor of Dr. Edward \tIcCracly. of the Uri 1`~1l`~' ` T
of the $outh. who first made a biological survey of i lllv y (yy,
Nllllllville Caves several years ago and discovered ther L`t`ll¥` ~
fauna. T _ V
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l tanooga. Tenn. y lm
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111- ` .21; 11.—.'74; p.——1.20; 0.-.60; 21.--.65; 5.-.82; c.—
1;. 1 .81; g.—.12. 1
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211- 11.—.23; 11.—.72; p.———1.18; 0.—.59; :1.—.6B; _ 51
` s.—.81;c.—.81. .
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I2 (imlogzbzrl Survey of 1-Ilrzlmnm
ly flared, very truncate tip. The p?11`m11€rCS are shortt 'll](l I p;u
are armed witI1 tI1ree, short, stout setae. (PI. II, fig. 5%) i ILIII
pu—
PS€ll(lZlll0l)ll[llHlll1llS Iodingi fluviatilis HGW subspeci a III
,_ . . ' CUI
I yjzr /om/1/y: Rock House Cave, I 1111Ie so1: ol .
· · ` ;_ 1IIIt
()ICHII(lCI`. I\1IarshaII County, Ala. . my
H0/otyjze nm/e: L.—4.35 111111.: I‘V.—I.49 111111.; ),— — ICI
1.00 111111.; w.——.34: d.—.23; h.—.68; p.—I.I€ _ 1~‘I1
.63: a.——.66; s.——.8I: c.—.84: g.—.I I. . iII`<`
.. . . . . . . . . . . . _ WI
Superlicially. _//nvm/zlzs 1S very S1Ill1IZl1` to }IIt(’I`?(l/ 11/11.
The })I`OIIOLlllII is narrower, tI1e elytral pubescence. t ugh .
extremely sI1ort and fine. is a little lIIOI`€ obvious az the [III
I median tooth of tI1e Iabr11111 is obsolete, but in other 1*; eos IIN
the two li()I`llIS see111 outwardly identical. The aedea x ol Q I""
I/zmizz/1/is, however, is considerably shorter LIIZIII 1 1 ul A lull
nzerir/ioxm/is, a11d its IIIC(IIZlI`1 lobe. viewed dorsally, eni i11·: _ °"’I
wider. more truncate process. (PI. II, fig. 5) » IIIV
. IIIII
.·\ S1IIg`IC male was collected by Dr, I~VaIter B. _]o » on s mi]
the 23rd of `IUIIC, I9¢I-2, i11 a cave but six 111iIes {rom tI1 t‘II· 1 Wl,
nessee River a11d Iiilteen miles 11ortI1west ol the type I 1I1t\’ ° CM
ol; }}I(’I`l(lI())I{l/IA`. The unique Find is an unusually darI— >Cl`1· I yu,
I men. bei11g Iierruginous with pale elytral apex and appei @@*1 ,;,,,
I 111iI
PSC}!(l£\lIOpll[hHlll1llS Iodingi distiiigueus HCW Sl1I>S}>C<` , {pim
Type /om/[ty: IlIgC Cave, 5%, miles south ol VI IIIIW IIN
Morgan County, Ala. . gm
_ Lllh
'liyjxe seiiesz Holotype, allotype a11d 4 paratypt YPU < IM
locality, Sept. I4 ISI47, _Iones and Royer. my,
Holofy/Jr nm/ez L.——¢l-.38 111111.; I1\I.—I.46 111111. I-f z
I.I)II 111111.: w.—-.33; d.—.23; h.—.66: p.—I.II BJ · s PW
.60: a.—.67: s.—.80; c.—.76; ` 1-
AUOf_)‘]2e fenzu/ez L.—4.#I»3 111111.: \1\I.—I.5O IIIIIIV We ,1
l.()() 111111.; w.—~.34: d.—.23; I1.—.68: p.—I.I ¤‘·‘” I
.6I: a.—.65; s.—.8'7; c.—.'73. ‘
Compared with other races of lodingi situated st I1 or §
the lennessee River. (ll.Yf2'}IgIl€7I.S` agrees in having rel web
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.\'ett~ Ana];/tt/za/nzitl Beetles front Ajyjm/ac/tiaiz Region I3
ttl pztt l-sided elytra, relatively wide pronotal base and rec-
It tan r hind angles, but differs from all so far considered in .
if pos ing a narrower, straighter-sided pronotum, the margin
ol w h. anterior to the hind angles, is interrupted by a low tii
ton ?ty or tubercle, the angle apices themselves being sharp
"l ._ ant newhat produced. Far more striking, however, are
T tht racters of the aedeagus. The median lobe of this organ
_ .' ter; ttes in a flared, truncate process the right contour of
e,. _ wh1 t widely rounded. The transfer apparatus is short and
arti tt in dorsal view and the setae of the parameres
_ T ver tall. (Pl. II, fig. 4) Ҥ
lt`, I it
gh . though rlistiizgttetis may demonstrate by these aberrant
xc trttj certain geographic isolation, it is easy to see how the
tt T rtttt ttld have evolved from an ancestor like ffttztiztltfés which
,t` pos s similar specializations in less exaggerated form. The
,{ lttttt ?n turn, is easily derivable from nzeridioimlis. ln this
{ J tor tion it is important to note that a line drawn between
· tht; ttions of these three races is a nearly straight one run-
i nin; trthwest—southeast parallel to the Tennessee five to ten
ln itiih tuth of it; and that, furthermore, each locality repre-
lj Wil separate, minor drainage area, the creek involved in
if [ Cite se heading north to the main river. Thus a very per-
l' S let; ial cline can doubtless be established between meri-
" tlitt ~‘ at the bend of the river and (ff.\`ff7IgI((’}IS. forty five
_ Util > the northwest, (or between the former and races t
` tart to the west departing still more drastically from type
tx A lllél t>es distingttens) when the caves of this interesting re-
3 QW Fold additional material. A race connecting [/ttitirtlilis
M illlt W/ingnens, now represented by a single teneral male gt
Vn llttl t intermediate locality, awaits description until it S€1`i€S
Q wt collected.
r · i· PSP .!lOI)htllZl]lllllS hubriehli IWW $P€€i€$
1 e`/Je fom/ity: Dougherty’s Cave. 4% miles north of ;
" gtt» Lebanon, Russell Co., Va.
j il`/JF Series: Holotype, allotype and l paratype. type
yi, , locality, july 25 1939, Hubricht.
E
l
14 Geological Survey of Alabama 0
Holotyjze ma/ez L.—3.80 mm.; \~\l.—l.24 mm. ).—
.73 mm.; w.——.33: d.———.l9; h.——.70; p.—l.l». t·,—~ V bas
.60; a.—.67; s.—.93; c.—l.O9; g.—.l4. (Pl. I, L gm f llnr
Al/otype frniale: L.—4.00 mm.; \lV.—l.30 min, ),— it
.80 mm.; w.—.3?>; d.—.20; 11.-.72; p.—l.l· ;_~,` ” my
.58: a.—.68; s.—.93; c.—.84. ‘
y l)liS(1RI1"l`I()N or VTYPE SERIES. Y apa
Body: Rather elongate, compressed; testaceou. very 2
linely alutaceous. , lhlt
Head: Normal, rather narrow, sides gently zi late; _‘ elyl
pale l`errugino—testaceous; microsculpture relatively use apii
r reticuloalutaceous: median tooth of labrum obsolete: 1 liaa i
tooth ol labium linely, shallowly emarginate. ` ml,
Pronotnnzz Normal, rather narrow, il//5tlis as t 1 .t< I Wit
wide, width at base 7//Stlis ol? width at apex; anterior lglty i mi;.
l rounded, sides gently sinuate along entire extent, p< vial — wu
angles sharply rectangular, basal (secondary) angles t· yell; ‘ Rn
obtuse, basal margin very slightly convex medially; lat l re · inf
llexed margin narrow; basal impressions rather deep. tl .nrv— r ern
ly reticulate; disc convex, median line deeply inn sul. snr
glabrous.
li/y/mz Normal, rather elongate, 5y,»“8ths as wide ; nn;. _ lie
Iinely, sparcely pubescent; humeri rounded, sides very illll cli:
arcuate. subparallel over anterior third, apices trunc , llz- l
hiscent; rellexed margin rather wide, conlluent with — ll~‘~7 _ Re
liovea at humeri, margin obsoletely serrate at humeri, Xian e
llirougliout: striae evident but weak. distinctly punctu i. _
curvetl portion ol? apical stria {iairly long, straight, cul L
ward proximally; humeral and medial sets ol margina apri-
lzle Widely distributed. especially the latter; humeral fllllll . ` ggj
, subrarinate, disc lilat. ·
t .·l/zjxmulzxgesz Normal, jaws rather short; subu limi i Evil
segment ol maxillary palpns Qths the length ol ternllt » W? »
l ment: second antennal segment approximately ll lll ` ll`· l’·°°
l length ol third. 0
Y Q lrfll
‘ 3
3
.
_ if T TTT +*" " "“ ‘``’ w1_ ..:2;
i Yew ;f)LOj)]lf/l(ll777,f£`l Beetles from Appalrzclzirnz Region 15 L
__ trleagzzsz l\‘Iedian lobe arciform, with basal bulb and [
.—- V bus gxure poorly defined; apical process short, tapering in
fil i (lor orofile to subtruncate, slightly flared apex; transfer
-. rip]; us composed of a shorter internal and a longer en-
T , yell g piece, the latter rounded at the tip; parameres rela-
` _ tirt ang, bearing three short setae. (Pl. I, fig. 4) .
‘ h iteworthy characters which set this unique species
li spa om others of the genus are the unusual discrepancy in *
M _ sizt ween the segments of the maxillary palpi, the near ;
ll i C(|ll · in length of the second and third antennal segments. I
I the ic separation of the medial sctigerous papillae of the
rc; _‘ eln margin and the peculiarly angular. truncate elytral
i. zlplt
“` _ problematic affinity, /rubric/zti seems to be either :1
` rtl; of an archaic stock or an extreme isolation product E
.t< I our ing from some such conservative ancestry as lzirsufus
ts ‘ Nil;. nrovide. The general body form and aedeagus are i
ui — su; ire of the latter species which inhabits caves of Powell p
it. · Rn lrainage, seventy miles or so to the southwest of
. intl fi. The waters from the neighborhood of Dough-
;v— t wi ne evidently join the Clinch River. These parallel [
tl. Wm are both tributaries of the Tennessee. S
’ l
i ‘e present species bears the name of its discoverer. Les- {
u. . llf “l`1Cll[, who made a biological survey of many Appala-
li dn ares in l939.
lr i f
l
.=,.: l,
ig — Re ices. {
` l
‘t· _ iq leannel, R.: L’Abeille; 35. {
»l leannel, R.; Archiv. Zoo]. Exp.; 71, 3; pp. ·lO3·500.
gs , ) _ l Valentine, l\I.: lonr. Elisha l\iIitchell Sc. Soc.; #16, 21 pp. {
·‘ . ‘ s
1 ¤ KN if Valentine, M.; lonr, Elisha Mitchell Sc. Soc.: ~l7, 2: pp. I
.1. » Ji.- ` `
if ‘ W 1 W7 Valentine, _]. M.; Jour. Elisha Mitchell Sc. Soc.; 53, 1: pp. i
A yy] W Valentine, ]. M.; Trang. Conn. Acad. Arts Scs.; $6; pp. I
· ‘~ , ` •
‘ 4
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Q .
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lf} (imlogicrzl Survey of Alabanm 1
1 1>1.~
EXl)lHllH[i0l1 0f plates. ‘
PLATE I i
1 Fig. 1. [).$`Cl((1(I)IO])]I1]I{l[}7'lU5 izzlcrmcdizzs Z'C7}1])l(3[()lI}· 11. ss; 11110- 1 `*
Lypc 1`c111z11c; iiiggill1)O[1lZlI]1’S Cave, )[n:Mi1111vi111· L‘11l1. ·
(18 X) .
Fig. 2. P. }II(I('}`{l([(’Ii 11. sp., holotypc 111:11c: Higgi111><>1l1;1111's (l X1:» »
AI1I1I]\'111€, TCIIII. (18 ,
Fig. 31. P. /ll/1J1`1(`/I/1 11. sp., 1]O101.yl)C 111:11c; DOllg`1]C1`l}1',S (1 11J·
1)2l1]Ol], Vu. (18 X) 1
Fig. ·1. ;\C(1CLlgllS 0f P. /11111110/1ti 11. sp., h010typc; (101`SZl1 view. 1*1 X1
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18 Grological Survey of Alabama V;
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PLAT1; Il °1~ `_
Acclcagi ol [)SCII([(l}I0])ll[]l{l[Il'll(.S` sps., dorsal views. _
Fig. l. P. I·}I[(fI`)II(f([lillS lcizzpleetoriii 11. ssp., holotypcz Higginl izIll`> 1
Cave, McNIi1111villc, Term. (95 X)
Fig. 2. P. marradczi 11. sp., paratypc; Higg`l]]bO[llH1ll’S Ca Nic- `
)Ii1111villc, TG1111. (95 X) `
Fig. fl. P. Iodfrzgi fullrrri, 11. comb., topotypc; TCIIIICSSCC ’ sl`I15. »
Cliattanooga, TG1111. (100 X) ·
Fig. ·l. P. lmlingi disiingzzczzs 11. ssp., holotypc; Inge Cave. iii}; .p
Ala. (l00 X) 1
Fig. 5. P. Iodizzgi fIu1»iatil1'.s` 11. ssp., holotypcg Rock House C. Ulv .
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